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To complete the proposed cycle, human 5-oxoprolinase (OPLAH), a 275 kDa homodimer, couples ATP hydrolysis to the opening of the ring structure to generate glutamate (Van der Werf et al., 1971

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Thiosulfate (S = +2) can be excreted into the plasma and or further oxidized to sulfite (S = +4) for sulfate-PAPS production (Fig